Most of the current lion population estimates are based on three different counting methods. The earliest method, used in the 1970s and 1980s, relied on luring lions with bait. Researchers would catch a lion, tranquilize it, brand it, and release it back to the wild. If they encountered the branded lion again, they would know it had already been counted.
Today, most researchers use either call-up surveys or track counts. To do a call-up survey, “you drive to strategic locations, and you play the calls of a buffalo or a hyena,” says Braczkowski. After hearing the sounds of their prey, “lions run in, and you count each individual.” For track counts, researchers look for lion paw prints on the ground and count each set of tracks they find. Each set of tracks identified is counted as one lion.
The problem with these methods, Braczkowski says, is that it’s very easy to count the same animal several times. There’s no way to know if the same lion comes to multiple call-up surveys, or if multiple sets of tracks belong to the same lion. Track counting is particularly unreliable, as lion tracks may stay visible for a long time in dry environments but vanish quickly after a rainstorm or in windy areas.
Braczkowski is also concerned with how the findings from these counting methods are expanded to estimate populations over large areas. “The bigger assumption these surveys make is that they sample a small area of 200 square kilometers and extrapolate from that out to 2,000 square kilometers,” he says.